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Get Received 7 February 2000 Received In Revised Form 1 June 2000 Accepted 14 June 2000

987; Struhsaker, 1997) or subject to edge e ects'' (Malcolm, 1994, 1997). Although it would be convenient to use studies of mammals in undisturbed and disturbed habitats as a proxy for comparing protected and unprotected areas, it is known that responses vary between species (Di endorfer et al., 1995; Songer et al., 1997; Hayward et al., 1999); responses depend on the length of time since the habitat was altered (Je rey, 1977; Ramirez and Hornocker, 1981) and on the type of disturbance invo.

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